IB Biology · Theme D · D1.1

The most
important
photocopier.

DNA makes copies of itself with astonishing accuracy — and the machinery to do it has been working since LUCA.

9Sub-topics
39Key terms
SL+HLLevel
MoleculesLevel of organisation
D1.1
Why this topic

What this topic answers.

Every sub-topic below feeds at least one of these questions.

Guiding question 1

How is new DNA produced?

Guiding question 2

How has knowledge of DNA replication enabled applications in biotechnology?

D1.1.1 – D1.1.5 · Standard Level

5 things to lock in.

The required syllabus content for D1.1, in order. Each card is one lesson-sized checkpoint.

D1.1.1

DNA replication as production of exact copies of DNA with identical base sequences

DNA replication as production of exact copies of DNA with identical base sequences

D1.1.2

Semi-conservative nature of DNA replication and role of complementary base pairing

Semi-conservative nature of DNA replication and role of complementary base pairing

D1.1.3

Role of helicase and DNA polymerase in DNA replication

Role of helicase and DNA polymerase in DNA replication

D1.1.4

Polymerase chain reaction and gel electrophoresis as tools for amplifying and separating DNA

Polymerase chain reaction and gel electrophoresis as tools for amplifying and separating DNA

D1.1.5

Applications of polymerase chain reaction and gel electrophoresis

Note: The sequence of subtopics have been changed for the SL section of this topic, to improve clarity.

D1.1.1 · DNA replication overview

One DNA → two identical copies.

Before any cell divides — for growth, tissue replacement, or reproduction — it must first copy its DNA precisely. Each daughter cell needs the full genome.

DNA replication produces two daughter molecules with sequences identical to the original. This precision is what allows genetic information to pass faithfully from cell to cell, generation to generation.

When does it happen? Before mitosis (for growth and tissue replacement) and before meiosis (for sexual reproduction). The S phase of the cell cycle.

D1.1.2 / D1.1.3 · Mechanism & semi-conservative

Semi-conservative replication, in three steps.

Each daughter helix has one old strand and one new — proven by Meselson & Stahl in 1958. Each strand serves as the template for its partner.

  1. Unwinding. Helicase breaks the hydrogen bonds between complementary base pairs, unwinding the double helix into two separate strands.
  2. Base-pairing. Each separated strand acts as a template. Free nucleotides pair with their complements: A↔T, G↔C.
  3. Polymerisation. DNA polymerase joins the new nucleotides into a growing strand, using the original as a template.

Result: two identical double helices, each with one parent strand and one new strand. Accuracy comes from the obligate complementary base pairing — no other combinations fit.

D1.1.4 · Polymerase chain reaction

Making billions of copies from a tiny sample.

PCR amplifies a chosen DNA sequence in vitro, using cycles of heating and cooling. The single technique that powers forensics, diagnostics, and most molecular biology.

The PCR cycle

  1. Denaturation (95 °C) — heat breaks hydrogen bonds between strands.
  2. Annealing (~54 °C) — temperature drops; primers bind to the start of the target sequence on each strand.
  3. Extension (72 °C) — Taq DNA polymerase extends from each primer, building a new complementary strand.
  4. Repeat — each cycle doubles the DNA. 30 cycles → over a billion copies.
🌋

Taq polymerase — heat-stable from hot springs

Normal DNA polymerases denature at 95 °C. The whole technique would be impractical if you had to add fresh enzyme every cycle. Taq polymerase from Thermus aquaticus (a bacterium that lives in Yellowstone's hot springs) is heat-stable up to ~98 °C. One addition lasts all 30 cycles. The breakthrough that made PCR practical.

D1.1.4 · Gel electrophoresis

Separating DNA by size.

DNA fragments cut by restriction enzymes can be separated by length using an electric field through an agarose gel.

  1. DNA is digested with restriction enzymes that cut at specific sequences, producing fragments of various sizes.
  2. Fragments are loaded into wells in an agarose gel placed in a salt solution.
  3. An electric current is applied. DNA is acidic and negatively charged — moves toward the positive electrode.
  4. Smaller fragments move through the gel faster than larger ones (less resistance).
  5. A dye reveals the pattern of bands — the DNA profile.
D1.1.5 · Applications

From crime scenes to paternity tests.

PCR + gel electrophoresis enables DNA profiling, used in forensics, paternity testing, evolution studies, disease diagnosis, and more.

Forensics

Matching crime-scene DNA

Tiny DNA samples from a crime scene (hair, blood, skin) are amplified by PCR, then profiled by gel electrophoresis. The pattern is compared with profiles from suspects. All bands must match for a confident identification.

Paternity testing

Whose child?

A child shares half of its DNA with each parent. Every band in the child's profile must appear in either the mother's or the father's profile. If they do, parentage is confirmed.

📊

Nature of Science · reliability through more markers

The reliability of a DNA profile depends on the number of markers used. More markers → smaller probability of a chance match → fewer false identifications. Modern forensic profiling uses 13+ markers.

HL extension

Higher Level only.

An extra 4 sub-topics for HL — same syllabus, deeper mechanism.

HL only

Directionality of DNA polymerases

Directionality of DNA polymerases

HL only

Differences between replication on the leading strand and the lagging strand

Differences between replication on the leading strand and the lagging strand

HL only

Functions of DNA primase, DNA polymerase I, DNA polymerase III and DNA ligase in replication

Functions of DNA primase, DNA polymerase I, DNA polymerase III and DNA ligase in replication

HL only

DNA proofreading

DNA proofreading

D1.1.6 · Directionality

DNA polymerase only goes one way.

DNA polymerases add new nucleotides only to the 3' end of an existing strand — they synthesise 5' → 3'. This constraint shapes everything about how replication works.

Each nucleotide's 5' phosphate attaches to the previous nucleotide's 3' hydroxyl — so the strand grows from the 5' end outward. Since the two parent strands are antiparallel (5'→3' one way; 3'→5' the other), the new strands are also antiparallel — and they get synthesised differently on the two sides of the replication fork.

D1.1.7 · Leading vs lagging strand

Continuous, or in fragments.

Because polymerase only adds in the 5' → 3' direction, one new strand can be made continuously and the other must be made in pieces.

Leading strand

Continuous synthesis

Synthesised in the same direction as the replication fork moves. Polymerase keeps adding nucleotides without interruption. Just one RNA primer needed at the start.

Lagging strand

Discontinuous synthesis

Synthesised away from the fork — only short stretches can be made before having to restart. Each stretch is an Okazaki fragment. Multiple primers are required — one for each fragment. Fragments are later joined by DNA ligase.

D1.1.8 · The replication enzymes

Five enzymes working together.

Prokaryotic DNA replication is the IB's reference case. Five enzymes do the job.

EnzymeFunction
HelicaseUnwinds the double helix at the replication fork by breaking H-bonds.
Gyrase (topoisomerase)Relieves the supercoiling strain that builds up ahead of the fork.
DNA primaseSynthesises short RNA primers to initiate replication.
DNA polymerase IIIAdds DNA nucleotides to the 3' end of each primer (5' → 3' direction). Main polymerase.
DNA polymerase IRemoves RNA primers and replaces them with DNA nucleotides.
DNA ligaseJoins Okazaki fragments on the lagging strand.
D1.1.9 · DNA proofreading

Catching errors as they happen.

DNA polymerase III doesn't just add nucleotides — it also reads back what it just added, and corrects mistakes on the spot.

After adding each nucleotide, DNA polymerase III checks for correct base pairing. If a mismatch is detected at the 3' terminal, the polymerase backs up, removes the mis-matched nucleotide, and replaces it with the correct one before resuming. This proofreading lowers the error rate from about 1 in 10⁴ to about 1 in 10⁶ — roughly 100× more accurate. Additional repair pathways after replication push the final error rate to about 1 in 10⁹.

HL-only key terms

DNA PrimaseRNA PrimerDNA Polymerase IIIDNA Polymerase IDNA LigaseGyraseReplication ForkLeading StrandLagging StrandContinuous ReplicationDiscontinuous ReplicationOkazaki FragmentsDNA ProofreadingDNA Replication
Vocabulary

25 terms to own.

If you can't define one of these in a sentence, that's where to revise next.

DNANucleotideDNA ReplicationHelicaseDNA PolymeraseComplementary Base PairsSemi-ConservativePolymerase Chain Reaction (PCR)PrimersTaq DNA PolymeraseDenaturationAnnealingGel ElectrophoresisDNA ProfileRestriction EndonucleaseDNA MarkersDiagram of DNADraw a labelled diagram of DNAStructure of DNALabelled Diagram of DNAPhosphateHydrogen BondsDeoxyriboseCovalent BondNitrogen Base

IB Linking Questions

“How is genetic continuity ensured between generations?”

“What biological mechanisms rely on directionality?”