IB Biology · Theme C · C4.2

Energy flows,
matter cycles.

Energy flows once and is lost as heat. Matter cycles forever. Two laws that shape every ecosystem on Earth.

13Sub-topics
0Key terms
SL+HLLevel
EcosystemsLevel of organisation
C4.2
Why this topic

What this topic answers.

Every sub-topic below feeds at least one of these questions.

Topic focus

Transfers of Energy and Matter.

C4.2.1 – C4.2.13 · Standard & Higher Level

13 things to lock in.

The required syllabus content for C4.2, in order. Each card is one lesson-sized checkpoint.

C4.2.1

Food chains and food webs

Food chains and food webs

C4.2.2

Trophic levels: producers, consumers, decomposers

Trophic levels: producers, consumers, decomposers

C4.2.3

Detritivores vs decomposers; saprotrophic nutrition

Detritivores vs decomposers; saprotrophic nutrition

C4.2.4

GPP, NPP, and the NPP = GPP − R equation

NPP = GPP − Respiration | Units: kJ m⁻² yr⁻¹ or g dry mass m⁻² yr⁻¹

C4.2.5

Energy transfer efficiency; the ~10% rule

Energy transfer efficiency; the ~10% rule

C4.2.6

Reasons for energy loss between trophic levels

Reasons for energy loss between trophic levels

C4.2.7

Ecological pyramids: numbers, biomass, energy

Count of organisms at each level.

C4.2.8

Carbon cycle: photosynthesis, respiration, decomposition

Carbon cycle: photosynthesis, respiration, decomposition

C4.2.9

Combustion, fossilisation; carbon sinks and sources

Combustion, fossilisation; carbon sinks and sources

C4.2.10

Saprotrophic nutrition — enzyme secretion and absorption

How Saprotrophic Nutrition Works

C4.2.11

Nitrogen cycle: fixation, nitrification, denitrification, ammonification ★HL

Nitrogen cycle: fixation, nitrification, denitrification, ammonification ★HL

C4.2.12

Leaching, eutrophication and soil fertility ★HL

Leaching, eutrophication and soil fertility ★HL

C4.2.13

Biomagnification and bioaccumulation (DDT) ★HL

IB Biology DP | C4.2 Transfers of Energy and Matter | SL & HL

C4.2.1 / C4.2.2 · Food chains, webs, trophic levels

Energy flows one way up the levels.

Food chains run from producers up through consumers. Food webs are networks of interconnected chains. Arrows show direction of energy flow, not predation.

LevelRoleExample
TL 1 · ProducerAutotroph — fixes CO₂ by photosynthesisOak tree
TL 2 · Primary consumerHerbivoreCaterpillar
TL 3 · Secondary consumerCarnivore eating herbivoresBlue tit
TL 4+ · Tertiary consumerApex predatorSparrowhawk
C4.2.3 · Detritivores vs decomposers

Two ways to recycle dead matter.

Both break down dead material — but differently. Together they're essential for nutrient cycling.

Detritivores

Animals — internal digestion

Earthworms, woodlice, dung beetles. Ingest dead matter; digest it inside their gut (intracellular). Fragment large pieces into smaller ones, increasing surface area for decomposers.

Decomposers

Bacteria, fungi — external digestion

Saprotrophic. Secrete enzymes onto dead matter (extracellular); absorb soluble products. Release mineral ions (NH₄⁺, PO₄³⁻) directly into the soil.

C4.2.4 · Primary productivity

GPP, NPP, and the difference.

Producers fix energy from sunlight. Some they use themselves for respiration; the remainder is available to consumers.

📐

The equation

NPP = GPP − R

GPP = gross primary productivity (total energy fixed). R = energy lost to respiration. NPP = net primary productivity = energy available to consumers. Units: kJ m⁻² yr⁻¹ or g dry mass m⁻² yr⁻¹.

Example: Forest GPP = 8,500 kJ m⁻² yr⁻¹; respiration = 3,200 kJ m⁻² yr⁻¹; NPP = 5,300 kJ m⁻² yr⁻¹. ~62% of GPP becomes available to consumers.

C4.2.5 / C4.2.6 · Energy transfer efficiency

Roughly 10% passes upward.

Only a small fraction of energy at one trophic level reaches the next. Most is lost as heat from respiration.

Typical efficiency
~10 %

Range 5–20% across ecosystems.

Heat from respiration
major

The biggest single loss. Lost forever to space.

Egestion
faeces

Undigested material never absorbed; goes to decomposers, not the next level.

Uneaten biomass
death

Organisms that die without being eaten — feed decomposers, not predators.

This is why food chains are short — by the 4th or 5th trophic level, there's barely any energy left to support large populations.

C4.2.7 · Ecological pyramids

Three ways to draw a food web.

Pyramids of numbers, biomass, and energy each tell a different story about an ecosystem.

Numbers

Count per level

Can be inverted — one large tree supports millions of insects. Least informative because organism size is ignored.

Biomass

Dry mass per level (g m⁻²)

Can be inverted in some systems — phytoplankton reproduce so fast that their standing biomass is less than zooplankton biomass at any instant.

Energy

Rate of energy flow (kJ m⁻² yr⁻¹)

Always upright. Energy must decrease at each level — second law of thermodynamics. The most accurate representation.

C4.2.8 / C4.2.9 · Carbon cycle

Carbon moves between organic and inorganic forms.

Four processes drive the carbon cycle. Two add CO₂ to the atmosphere; two remove it. Plus geological-timescale sinks.

Photosynthesis

CO₂ → organic carbon

Producers fix atmospheric CO₂ into glucose using light energy. Major removal pathway. Performed by plants, algae, cyanobacteria.

Cellular respiration

Organic carbon → CO₂

All organisms release CO₂ as they oxidise glucose for ATP. Continuous, day and night.

Decomposition

Dead matter → CO₂

Saprotrophs break down dead organic matter, releasing CO₂ and mineral nutrients. The recycling channel.

Combustion

Fossil/biomass → CO₂

Burning fossil fuels (coal, oil, gas) releases carbon fixed over millions of years. The major anthropogenic driver of rising atmospheric CO₂.

Fossilisation & sinks

Dead organisms buried under heat and pressure for millions of years → coal, oil, natural gas. A geologically slow carbon sink. Oceans, forests and peat bogs are biological carbon sinks — they absorb more CO₂ than they release on contemporary timescales.

Vocabulary

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