IB Biology · Theme B · B3.3 · HL only

How
movement
happens.

Filaments slide past filaments. ATP burns. Bodies move. Cells, sarcomeres, joints.

10Sub-topics
56Key terms
HL onlyLevel
OrganismsLevel of organisation
B3.3
Why this topic

What this topic answers.

Every sub-topic below feeds at least one of these questions.

Guiding question 1

How do muscles contract and cause movement?

Guiding question 2

What are the benefits to animals of having muscle tissue?

HL extension

Higher Level only.

An extra 10 sub-topics for HL — same syllabus, deeper mechanism.

HL only

Adaptations for movement as a universal feature of living organisms

Adaptations for movement as a universal feature of living organisms

HL only

Sliding filament model of muscle contraction

Sliding filament model of muscle contraction

HL only

Role of the protein titin and antagonistic muscles in muscle relaxation

Role of the protein titin and antagonistic muscles in muscle relaxation

HL only

Structure and function of motor units in skeletal muscle

Structure and function of motor units in skeletal muscle

HL only

Roles of skeletons as anchorage for muscles and as levers

Roles of skeletons as anchorage for muscles and as levers

HL only

Movement at a synovial joint

Movement at a synovial joint

HL only

Range of motion of a joint

Range of motion of a joint

HL only

Internal and external intercostal muscles as an example of antagonistic muscle action to facilitate internal body movements

Internal and external intercostal muscles as an example of antagonistic muscle action to facilitate internal body movements

HL only

Reasons for locomotion

Reasons for locomotion

HL only

Adaptations for swimming in marine mammals

Adaptations for swimming in marine mammals

B3.3.1 · Movement everywhere

Every organism moves something.

From bacteria following a sugar gradient to humans running marathons, movement is a universal feature of living things. Even "sessile" organisms move parts of themselves.

Motile

Whole organism moves

Bacteria use flagella to swim toward food.
Amoeba moves by reorganising its cytoskeleton, extending pseudopodia.
Paramecium swims using thousands of cilia.
Mammals use muscles attached to bones.

Sessile

Anchored, but parts move

Adult barnacles are attached to rocks but extend feathery cirri to filter food from water.
Plants show tropisms — directional growth toward stimuli like light (phototropism).

B3.3.2 · Sliding filament model

Filaments slide. Sarcomeres shorten.

Muscle contraction is the sliding of actin and myosin filaments past each other within sarcomeres — not the shortening of the filaments themselves.

The structure

  • Sarcolemma — plasma membrane of the muscle fibre.
  • Sarcoplasm — its cytoplasm.
  • Sarcoplasmic reticulum — modified ER, stores Ca²⁺.
  • Mitochondria — abundant; supply ATP for contraction.
  • Myofibrils — contractile filaments running along the fibre.
  • Sarcomeres — repeating units of each myofibril, defined by Z lines at each end, with thin actin filaments anchored at the Z lines and thick myosin filaments centred on the M line.

The contraction sequence

  1. An action potential arrives at the muscle fibre via a motor neuron.
  2. The action potential stimulates the sarcoplasmic reticulum to release Ca²⁺.
  3. Ca²⁺ binds to troponin on actin filaments → troponin changes shape.
  4. Troponin shifts tropomyosin, exposing the myosin binding sites on actin.
  5. Myosin heads bind to actin, forming cross-bridges.
  6. ATP binds to myosin head → cross-bridge breaks.
  7. ATP is hydrolysed to ADP + Pi → myosin head cocks (moves toward the Z line).
  8. ADP is released → power stroke: myosin pulls actin toward the centre of the sarcomere.
  9. Cycle repeats. Sarcomeres shorten; muscle contracts.
B3.3.3 · Titin and antagonistic muscles

Muscles can't push.

Muscle tissue only exerts force by contracting. Returning to its relaxed state needs something else — either a molecular spring (titin) or an opposing muscle (antagonist).

Titin

The molecular spring

Titin spans from the Z line to the M line through each sarcomere. It is highly elastic — helps the sarcomere recoil after stretching, and prevents overstretching during force production.

Antagonistic pairs

One contracts, the other stretches

Muscles work in opposing pairs. When biceps contracts (flexing elbow), triceps relaxes and stretches. To extend the elbow, triceps contracts and biceps relaxes.

B3.3.4 · Motor units

From brain to fibre.

A motor unit is one motor neuron plus all the muscle fibres it innervates. The neuromuscular junction is where the signal jumps from nerve to muscle.

Motor neurons can be over 1 m long in humans (and over 5 m in giraffes). Their structure:

  • Dendrites — receive input from other neurons.
  • Cell body — integrates signals, decides whether to fire.
  • Axon — long projection conducting action potentials, wrapped in Schwann cells forming myelin.
  • Axon terminals — release neurotransmitter (acetylcholine) onto the muscle fibre.

At the neuromuscular junction, acetylcholine released by the motor neuron binds to receptors on the sarcolemma, triggering an action potential in the muscle fibre.

B3.3.5 · Skeletons

Anchors and levers.

Skeletons give muscles something to pull against, and translate muscle contraction into useful body movement.

Vertebrates

Endoskeleton

Internal skeleton. Bones are inside; muscles attach to the outside of bones via tendons. Bones act as levers — contraction of muscles produces movement around joints.

Arthropods

Exoskeleton

External skeleton made of chitin. Muscles attach to the inside of the exoskeleton. Same lever principle — the exoskeleton transfers force into movement.

B3.3.6 · The synovial joint

Six parts working together.

A synovial joint is a freely movable joint with a fluid-filled capsule. The IB-named example is the hip — femur fitting into the acetabulum of the pelvis.

StructureFunction
BonesAct as levers. Femur + pelvis in the hip joint.
CartilageCovers bone ends — reduces friction, absorbs shock.
Synovial fluidLubricates cartilage; supplies nutrients and O₂ to cartilage.
Joint capsuleSeals the synovial fluid in; limits joint movement.
TendonsAnchor muscles to bones.
LigamentsConnect bones to bones; stabilise the joint.
B3.3.7 · Range of motion

Hinge or ball-and-socket.

Two joint types with different movement capabilities. Range of motion can be quantified with a goniometer.

Hinge joint

One plane · flexion + extension

Allows movement in only one direction (like a door hinge). Examples: elbow, knee, finger joints. Flexion (bending) and extension (straightening).

Ball-and-socket

Multiple planes · flexion + extension + rotation

Spherical bone head fits into cup-shaped socket. Allows flexion, extension, abduction, adduction, and rotation. Examples: hip, shoulder.

Goniometry measures joint angles with a goniometer. Used in physiotherapy to assess injuries, recovery, and to compare healthy ranges of motion.

B3.3.8 · Antagonistic intercostals

The ribcage's internal antagonists.

Internal and external intercostal muscles are oriented in opposite diagonal directions — when one contracts it stretches the other, storing energy in titin.

  • External intercostals — closer to the surface, oriented parallel to the muscle. Contract → lift ribcage up and out (inspiration).
  • Internal intercostals — deeper, oriented diagonally. Contract → pull ribcage down and in (forced expiration).

Their opposite orientations mean each layer stretches the other when it contracts — a textbook example of antagonistic muscle action in internal body movement.

B3.3.9 · Reasons for locomotion

Why animals go places.

Four major drivers of locomotion in animals — each with named examples.

1

Foraging

Moving to find food. From grazing herbivores to predators tracking prey, foraging is the most universal driver.

2

Escape

Moving away from threats. Pronghorns have evolved extreme speed to outrun (now-extinct) predators.

3

Finding a mate

Moving to encounter or attract mates. Male birds of paradise perform elaborate dance routines to attract females.

4

Migration

Seasonal long-distance movement for food, breeding or climate. Wildebeest follow rainfall across the Serengeti.

B3.3.10 · Marine mammal adaptations

Land mammals, re-engineered for water.

Whales, dolphins and seals evolved from terrestrial ancestors. Their adaptations to swimming illustrate evolutionary plasticity.

  • Streamlined body — minimises drag while swimming.
  • Modified pentadactyl limbs — flippers in seals and dolphins, evolved from the same five-fingered limb plan as our hands.
  • Fluke (modified tail) — flat horizontal tail in whales and dolphins. Moves up and down, powered by huge muscles.
  • Modified airways — blowholes are modified nostrils on top of the head, allowing breathing while the body stays submerged.
  • Large lung capacity — stores oxygen for long dives.
  • Muscular nostril control — prevents water entering during dives.
  • Rigid bronchial tubes — prevent lung collapse under deep-water pressure.

HL-only key terms

MovementMotileSessileMuscle fibresSarcolemmaSarcoplasmSarcoplasmic ReticulumMyofibrilsSarcomeresDark BandLight BandZ linesM linesTroponinTropomyosinPowerstrokeSliding Filament TheoryActinMyosinTitinAntagonistic MusclesMotor NeuronsAction PotentialsDendritesCell BodyAxonSchwann CellsAxon TerminalsNeuromuscular JunctionSynapseAcetylcholineJointSynovial JointTendonsLigamentsCartilageSynovial FluidJoint CapsuleExoskeletonsBall and Socket JointHinge JointFlexionExtensionRotational MotionGoniometryGoniometerLocomotionForagingMigrationMammalEndothermicMarine MammalsPentadactyl LimbFluke (modified tail)EndoskeletonExoskeleton
Vocabulary

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IB Linking Questions

“What are the advantages and disadvantages of dispersal of offspring from their parents?”

“In what ways does locomotion contribute to evolution within living organisms?”