IB Biology · Theme B · B3.2

The arteries
of every
large body.

Diffusion gets you only so far. Beyond about a millimetre, you need a circulation — and an engine to drive it.

18Sub-topics
46Key terms
SL+HLLevel
OrganismsLevel of organisation
B3.2
Why this topic

What this topic answers.

Every sub-topic below feeds at least one of these questions.

Guiding question 1

What adaptations facilitate transport of fluids in animals and plants?

Guiding question 2

What are the differences and similarities between transport in animals and plants?

B3.2.1 – B3.2.10 · Standard Level

10 things to lock in.

The required syllabus content for B3.2, in order. Each card is one lesson-sized checkpoint.

B3.2.1

Adaptations of capillaries for exchange of materials between blood and the internal or external environment

Adaptations should include a large surface area due to branching and narrow diameters, thin walls, and fenestrations in some capillaries where exchange needs to be particularly rapid.

B3.2.2

Structure of arteries and veins

Structure of arteries and veins

B3.2.3

Adaptations of arteries for the transport of blood away from the heart

Adaptations of arteries for the transport of blood away from the heart

B3.2.4

Measurement of pulse rates

Measurement of pulse rates

B3.2.5

Adaptations of veins for the return of blood to the heart

Adaptations of veins for the return of blood to the heart

B3.2.6

Causes and consequences of occlusion of the coronary arteries

Causes and consequences of occlusion of the coronary arteries

B3.2.7

Transport of water from roots to leaves during transpiration

Transport of water from roots to leaves during transpiration

B3.2.8

Adaptations of xylem vessels for transport of water

Adaptations of xylem vessels for transport of water

B3.2.9

Distribution of tissues in a transverse section of the stem of a dicotyledonous plant

Distribution of tissues in a transverse section of the stem of a dicotyledonous plant

B3.2.10

Distribution of tissues in a transverse section of the root of a dicotyledonous plant

Distribution of tissues in a transverse section of the root of a dicotyledonous plant

B3.2.1 · Capillaries

Where the actual exchange happens.

Arteries deliver. Veins return. Capillaries are where blood actually exchanges materials with the tissues — and their structure is exquisitely adapted for that one job.

Large surface area

Branching dense networks

Capillaries are densely branched throughout tissues, providing huge total surface area for exchange.

Thin walls

One cell thick

Walls are a single layer of squamous endothelial cells — minimum possible diffusion distance.

Narrow lumen

Single-file traffic

Lumen just wide enough for one red blood cell at a time — slows flow and maximises exchange time. Some capillaries also have fenestrations (small pores) for even faster exchange.

B3.2.2 / B3.2.3 · Arteries

Built to handle high pressure.

Arteries carry blood away from the heart at high pressure. Their wall structure has evolved to withstand and even smooth that pressure.

In a micrograph, you can tell an artery from a vein at a glance: arteries have relatively thick walls and narrow lumens; veins have thin walls and wide lumens.

B3.2.4 · Pulse

Feel the artery wall recoil.

Each heartbeat sends a pressure wave through the arteries — the pulse. You can feel it where an artery runs close to the skin.

Modern oximeters and smartwatches measure pulse optically (photoplethysmography). Worth comparing your manual count to a digital reading — both have failure modes.

B3.2.5 · Veins

Returning blood against gravity.

By the time blood reaches the veins, it has lost most of its pressure. Veins are adapted to keep it moving anyway.

B3.2.6 · Coronary heart disease

When the heart's own supply fails.

Coronary arteries feed the heart muscle itself. Their occlusion by atherosclerotic plaque causes heart attacks — and most of the risk factors are modifiable.

Atherosclerosis develops in stages:

  1. The inner artery wall is damaged (often by high blood pressure or smoking).
  2. Macrophages migrate to the damaged site, consume cholesterol from blood, and start forming a fatty streak.
  3. Fibrous tissue grows around the streak — a plaque forms.
  4. Plaque accumulates over years, narrowing the lumen.
  5. Plaque may rupture, triggering a blood clot that completely blocks the artery → heart attack.
Non-modifiable

Genetics · age · sex

Some genes increase risk. Older people have more damaged arteries. Males have higher risk on average.

Modifiable

Obesity · inactivity · smoking · diet

Obesity raises blood pressure and damages walls. Inactivity contributes to obesity. Smoking raises blood pressure and damages endothelium directly. High saturated fat / cholesterol diets contribute to plaque formation.

📊

Nature of Science · correlation vs causation

The Seven Countries Study showed a correlation between saturated fat intake and coronary heart disease deaths. Correlation coefficients quantify this — but correlation alone never proves causation. Many additional lines of evidence (mechanisms, randomised trials) support the causal link.

B3.2.7 · Transpiration pull

Water pulled up under tension.

The cohesion-tension theory in five steps. The driving force isn't push — it's pull.

  1. Water enters root cells by osmosis.
  2. Water moves into xylem in the roots.
  3. In leaves, water evaporates from mesophyll cells and diffuses out through stomata (transpiration).
  4. Water is drawn out of xylem through cell walls by capillary action (adhesion to cellulose) to replace what evaporated.
  5. Loss of water in the xylem creates negative pressure (tension) inside the xylem. The cohesive nature of water (hydrogen bonding) holds the column together; tension transmits downward all the way to the roots, pulling more water up.

This is the transpiration pull. It's strong enough to raise water more than 100 m to the canopy of a redwood tree.

B3.2.8 · Xylem adaptations

Dead cells, perfect plumbing.

Xylem vessels are highly specialised — they don't even have to be alive to do their job.

B3.2.9 / B3.2.10 · Stem and root anatomy

Where the tissues sit in plants.

In a transverse section of a dicotyledonous plant, vascular tissue (xylem + phloem) is arranged differently in stem vs root — and you should be able to draw a plan diagram of both.

Stem

Vascular bundles in a ring

Working from outside in: epidermis (protection) → cortex (support, starch storage) → vascular bundles arranged in a ring. Within each bundle, xylem is on the inside and phloem on the outside.

Root

Central vascular cylinder

Working from outside in: epidermis (often with root hairs for absorption) → cortex (broad layer for storage and water passage) → central vascular cylinder with xylem in a star or X pattern surrounded by phloem.

HL extension

Higher Level only.

An extra 8 sub-topics for HL — same syllabus, deeper mechanism.

HL only

Release and reuptake of tissue fluid in capillaries

Release and reuptake of tissue fluid in capillaries

HL only

Exchange of substances between tissue fluid and cells in tissues

Exchange of substances between tissue fluid and cells in tissues

HL only

Drainage of excess tissue fluid into lymph ducts

Drainage of excess tissue fluid into lymph ducts

HL only

Differences between the single circulation of bony fish and the double circulation of mammals

Differences between the single circulation of bony fish and the double circulation of mammals

HL only

Adaptations of the mammalian heart for delivering pressurized blood to the arteries

Adaptations of the mammalian heart for delivering pressurized blood to the arteries

HL only

Stages in the cardiac cycle

Stages in the cardiac cycle

HL only

Generation of root pressure in xylem vessels by active transport of mineral ions

Generation of root pressure in xylem vessels by active transport of mineral ions

HL only

Adaptations of phloem sieve tubes and companion cells for translocation of sap

Adaptations of phloem sieve tubes and companion cells for translocation of sap

B3.2.11–13 · Tissue fluid & lymph

Capillaries leak on purpose.

At the arterial end, fluid is forced out of capillaries into the tissues. At the venous end, most is drawn back in. The excess is collected by lymph vessels.

At the arterial end

Hydrostatic pressure wins

High blood pressure (hydrostatic) pushes fluid out through the thin capillary walls into the tissue spaces. The fluid (lacking large proteins) becomes tissue fluid bathing the cells.

At the venous end

Oncotic pressure wins

Blood pressure has fallen. Plasma proteins (mainly albumin) still inside the capillary draw water back in by osmosis — oncotic pressure. Most tissue fluid returns to the blood here.

In tissue fluid, substances exchange between blood and cells: O₂, CO₂, glucose, amino acids, ions, hormones — all diffuse or are actively transported between capillaries and the surrounding cells.

The fluid that doesn't return to capillaries (~10%) drains into lymph ducts. Lymph is returned to the bloodstream via the thoracic duct, which empties into a large vein near the heart.

B3.2.14 · Single vs double circulation

One trip or two.

Fish have single circulation — blood passes through the heart once per circuit. Mammals (and birds) have double circulation — blood passes through twice.

Single (fish)

Heart → gills → body → heart

Blood is pumped from the heart to the gills (gas exchange) and then directly on to the body, before returning to the heart. Pressure drops dramatically through the gill capillaries, so systemic flow is relatively slow.

Double (mammal)

Two pumps in one heart

Right side of heart pumps deoxygenated blood to the lungs (pulmonary circuit). Oxygenated blood returns to the left side, which pumps it to the body (systemic circuit) at high pressure. Each circuit gets full pump pressure — efficient systemic delivery to active metabolism.

B3.2.15 · Mammalian heart adaptations

Four chambers, two pumps, three valve sets.

The mammalian heart is two pumps in one organ — and is precisely engineered to deliver pressurised blood to the arteries.

  • Four chambers — left and right atria (upper), left and right ventricles (lower).
  • Septum separates left from right — keeps oxygenated and deoxygenated blood entirely separate.
  • Atrioventricular valves (tricuspid right, bicuspid/mitral left) prevent backflow into atria during ventricular contraction.
  • Semilunar valves (pulmonary and aortic) prevent backflow into ventricles during diastole.
  • Thick left ventricle wall — much thicker than the right, because it pumps blood to the entire body at higher pressure. Right ventricle pumps only to the lungs.
B3.2.16 · Cardiac cycle

One heartbeat in three phases.

The cardiac cycle is the repeating sequence of contractions and relaxations that constitutes a heartbeat. It is myogenic — generated by the heart itself.

  1. Atrial systole — atria contract, pushing remaining blood into ventricles. AV valves still open.
  2. Ventricular systole — ventricles contract. AV valves close (creating the first heart sound). Pressure rises until semilunar valves open and blood is forced into the aorta (left) and pulmonary artery (right).
  3. Diastole — all chambers relax. Semilunar valves close (second heart sound). Blood flows passively from veins into the atria, then into ventricles via the open AV valves. Cycle repeats.

Pacing is set by the sinoatrial (SA) node in the right atrium — the heart's intrinsic pacemaker. Impulses spread through the atria, pause at the AV node, then race down the bundle of His and Purkinje fibres to trigger ventricular contraction.

B3.2.17 · Root pressure

Pushing water up from below.

In addition to being pulled from above by transpiration, water in the xylem is sometimes pushed from below by root pressure.

In root endodermis cells, mineral ions are actively transported into the xylem against their concentration gradient. The resulting high solute concentration in xylem sap draws water in by osmosis — creating positive pressure that pushes water upward.

Root pressure is most evident at night when transpiration is low — visible as guttation (water droplets on leaf tips) in many plants. By day, the much greater transpiration pull dominates.

B3.2.18 · Phloem translocation

Sugars move from source to sink.

Phloem transports dissolved organic compounds (mainly sucrose, also amino acids) from places where they're made (sources) to places where they're used or stored (sinks).

  • Sources — mature photosynthesising leaves; storage organs releasing reserves (e.g. germinating seeds).
  • Sinks — growing tissues (apical and root meristems), fruits, storage organs being filled.
  • Sieve tube elements — phloem cells lined up end-to-end with perforated sieve plates between them. They lack nuclei and most organelles — built to be open conduits.
  • Companion cells — adjacent living cells, connected to sieve tubes by plasmodesmata. They have dense cytoplasm with many mitochondria. They provide the ATP needed for active loading of sucrose into the sieve tubes (phloem loading).

Mass flow: sucrose loaded at the source raises solute concentration → water enters by osmosis → high pressure pushes sap through sieve tubes toward sinks → at the sink, sucrose is unloaded; water follows back out. Pressure-driven mass flow.

HL-only key terms

Tissue FluidBlood PlasmaHydrostatic PressureOncotic PressureLymphLymph DuctsSingle CirculationDouble CirculationAtria/AtriumVentriclesAtrioventricular ValvesSemilunar ValvesCardiac cycleSystoleDiastoleMyogenicCasparian StripSieve TubesCompanion CellsPlasmodesmataTranslocationSourceSinkPhloem LoadingSinoatrial NodeAtrioventricular NodeBundle of HisPurkinje FibresAction PotentialRoot PressureSymplastic PathwayApoplastic Pathway
Vocabulary

14 terms to own.

If you can't define one of these in a sentence, that's where to revise next.

CapillaryArteryVeinLumenPulseCorrelation CoefficientsCoronary Heart DiseaseAtherosclerosisTranspirationXylem VesselsPhloemVascular TissueCortexEpidermis

IB Linking Questions

“How do pressure differences contribute to the movement of materials in an organism?”

“What processes happen in cycles at each level of biological organization?”