IB Biology · Theme B · B2.2

A cell
with rooms.

Eukaryotic cells aren't just bags of cytoplasm. They are factories with rooms — and a reason for the wall between each.

9Sub-topics
33Key terms
SL+HLLevel
CellsLevel of organisation
B2.2
Why this topic

What this topic answers.

Every sub-topic below feeds at least one of these questions.

Guiding question 1

How are organelles in cells adapted to their functions?

Guiding question 2

What are the advantages of compartmentalisation in cells?

B2.2.1 – B2.2.3 · Standard Level

3 things to lock in.

The required syllabus content for B2.2, in order. Each card is one lesson-sized checkpoint.

B2.2.1

Organelles as discrete subunits of cells that are adapted to perform specific functions

Nature of Science: Students should recognize that progress in science often follows development of new techniques.

B2.2.2

Advantage of the separation of the nucleus and cytoplasm into separate compartments

In prokaryotes this is not possible—mRNA may immediately meet ribosomes.

B2.2.3

Advantages of compartmentalisation in the cytoplasm of cells

Advantages of compartmentalisation in the cytoplasm of cells

B2.2.1 · What counts as an organelle

Discrete functional subunits.

Organelles are compartmentalised cellular structures with specific functions — and the IB has a particular list of what does and doesn't count.

Are organelles

Nucleus · vesicles · ribosomes · plasma membrane

Per the IB list. Each is a discrete structural unit with a defined function. (The plasma membrane is included by the IB, though not by most cell biologists, because it does compartmentalise.)

Not organelles

Cell wall · cytoplasm · cytoskeleton

Cell wall is extracellular. Cytoplasm is the bulk fluid, not compartmentalised. Cytoskeleton is dispersed throughout the cell, not bounded.

🔬

Nature of Science · technique drives progress

The function of individual organelles became studyable only when ultracentrifuges and cell fractionation techniques were developed in the 20th century. Each could now be isolated and its biochemistry examined separately.

Cell fractionation in three steps

  1. Homogenisation — blend tissue in a cold isotonic buffered solution to keep organelles intact.
  2. Filtration — remove larger cell debris.
  3. Ultracentrifugation — spin at increasing speeds. Densest organelles (nuclei) pellet first; lighter ones (mitochondria, ER, ribosomes) at higher speeds. Each pellet can be studied independently.
B2.2.2 · Nucleus / cytoplasm separation

Why the nuclear envelope matters.

Separating transcription from translation allows post-transcriptional modification of mRNA — something prokaryotes can't do.

In eukaryotes, transcription happens inside the nucleus. mRNA is then post-transcriptionally modified (capping, splicing, polyadenylation) before being exported through nuclear pores to the cytoplasm, where ribosomes translate it. This separation:

In prokaryotes there is no nucleus. mRNA is translated by ribosomes as it is being transcribed — no opportunity for post-transcriptional editing.

B2.2.3 · Cytoplasmic compartmentalisation

Why organelles are worth the cost.

Compartmentalising the cytoplasm enables concentrations, conditions and reactions that would be impossible in a single mixed pool.

Example 1

Lysosomes

Membrane-bound organelles full of digestive enzymes. The enzymes would shred the cell if released — the membrane keeps them in. When a cell dies, lysosomes burst and the enzymes self-digest the cell (autolysis).

Example 2

Phagocytic vacuoles

White blood cells engulf bacteria by phagocytosis, forming a phagocytic vacuole. Lysosomes fuse with the vacuole, releasing their enzymes to digest the bacterium. Compartmentalisation makes the destruction safe for the host cell.

HL extension

Higher Level only.

An extra 6 sub-topics for HL — same syllabus, deeper mechanism.

HL only

Adaptations of the mitochondrion for production of ATP by aerobic cell respiration

Adaptations of the mitochondrion for production of ATP by aerobic cell respiration

HL only

Adaptations of the chloroplast for photosynthesis

Adaptations of the chloroplast for photosynthesis

HL only

Functional benefits of the double membrane of the nucleus

Functional benefits of the double membrane of the nucleus

HL only

Structure and function of free ribosomes and of the rough endoplasmic reticulum

Structure and function of free ribosomes and of the rough endoplasmic reticulum

HL only

Structure and function of the Golgi apparatus

Structure and function of the Golgi apparatus

HL only

Structure and function of vesicles in cells

Structure and function of vesicles in cells

B2.2.4 · Mitochondrion adaptations

An organelle built for ATP.

Every part of the mitochondrion's structure serves the chemistry of aerobic respiration.

Outer membrane

Selective gateway

Protein channels let pyruvate in. Impermeable to H⁺ — allowing protons to accumulate in the intermembrane space.

Intermembrane space

Tiny — for fast H⁺ buildup

The narrow space between membranes means even small numbers of protons rapidly create a large gradient.

Inner membrane

Folded into cristae

Holds the electron transport chain and ATP synthase. Cristae fold dramatically increase surface area for these protein complexes.

Matrix

Krebs cycle factory

Contains the enzymes of the link reaction and Krebs cycle, plus mitochondrial DNA and 70S ribosomes that synthesise some respiratory proteins.

B2.2.5 · Chloroplast adaptations

An organelle built for photosynthesis.

Same logic as mitochondria — every structural feature serves the chemistry, this time splitting water and fixing carbon.

Thylakoids

Stacked discs (grana)

Flattened membrane discs containing chlorophyll, electron transport chain, and ATP synthase. Stacking into grana massively increases surface area for light absorption.

Thylakoid space

Tiny — for H⁺ accumulation

Like the mitochondrial intermembrane space — a small volume allows rapid proton gradient buildup.

Stroma

Calvin cycle factory

The fluid surrounding the thylakoids. Contains chloroplast DNA, 70S ribosomes, and all the enzymes of the Calvin cycle (light-independent reactions).

Double membrane

Selective gateway

Controls movement of CO₂ in and O₂ out; isolates chloroplast biochemistry from the cytoplasm.

B2.2.6 · The double nuclear membrane

Two functions for the nuclear envelope.

Why two membranes instead of one? It enables selective transport and graceful disassembly during cell division.

  • Nuclear pores are formed using integral proteins through both membranes. They allow selective passage of large molecules (mRNA out, proteins in). Single membranes can't support such large defined pores — they'd just leak.
  • Disassembly into vesicles at the start of mitosis/meiosis. The double membrane fragments into many small vesicles, allowing chromosomes to separate. At the end of division, vesicles fuse around the new daughter nuclei to rebuild the envelope.
B2.2.7 · Ribosomes & rough ER

Two ribosome addresses, two destinations.

The location of a ribosome determines where its protein ends up — keep-it-in-the-cell vs ship-it-out.

Ribosomes have two subunits (small and large), each made of rRNA and proteins. The small subunit binds mRNA; the large subunit catalyses peptide bond formation and has the polypeptide exit tunnel.

Free ribosomes

Cytoplasmic proteins

Float freely in the cytoplasm. Synthesise proteins that stay in the cell — glycolysis enzymes, cytoskeletal proteins, housekeeping enzymes — or that enter the nucleus.

Membrane-bound ribosomes

Export & lysosomal proteins

Attached to the rough ER. Synthesise proteins for secretion, for insertion into membranes, or for lysosomes. Proteins enter the ER lumen, get packaged into vesicles, and travel to the Golgi.

B2.2.8 · The Golgi apparatus

The post office of the cell.

Vesicles from the rough ER arrive at the Golgi, get modified and sorted, and leave as secretory vesicles bound for a specific destination.

The Golgi apparatus is a stack of flattened membrane sacs (cisternae). It sits between the rough ER and the plasma membrane.

  • Vesicles from rough ER bring proteins to the Golgi's cis face.
  • Enzymes inside the cisternae modify proteins — adding carbohydrates (making glycoproteins), or phosphate or sulfate groups; assembling quaternary structures.
  • Modified proteins exit the Golgi's trans face in secretory vesicles, bound for the plasma membrane (for exocytosis), lysosomes, or other destinations.

Two competing models describe how proteins move through the stack: the vesicle transport model (cisternae stay put; vesicles ferry proteins between them) and the cisternal maturation model (whole cisternae move from cis to trans, transforming as they go).

B2.2.9 · Vesicles & clathrin

Transport within the cell.

Vesicles are small phospholipid-bilayer sacs that carry cargo around the cell. Specific vesicles are built with the help of clathrin.

Vesicles transport proteins from the rough ER to the Golgi, between Golgi cisternae, and from the Golgi to the plasma membrane or to lysosomes. They also form during endocytosis to bring material into the cell.

Clathrin is a triskelion-shaped protein (three legs splayed out from a centre). When recruited to a region of membrane, clathrin proteins polymerise into a basket-like cage around that region, forcing the membrane to curve inward and bud off as a vesicle. The clathrin cage then disassembles, leaving a free vesicle ready to deliver its cargo.

HL-only key terms

MitochondrionAerobic RespirationIntermembrane SpaceCristaeMatrixKrebs CycleChloroplastPhotosynthesisThylakoidCalvin CycleStromaGranumRibosomesLarge Ribosomal SubunitSmall Ribosomal SubunitRough Endoplasmic ReticulumGolgi ApparatusVesicleClathrin
Vocabulary

14 terms to own.

If you can't define one of these in a sentence, that's where to revise next.

OrganellesCell FractionationUltrafiltrationPlasma MembraneHomogenizationNucleusCytoplasmTranscriptionTranslationMetabolitesEnzymesLysosomePhagocytosisPhagocytic Vacuole

IB Linking Questions

“What are examples of structure–function correlations at each level of biological organization?”

“What separation techniques are used by biologists?”