IB Biology · Theme C · C2.2

The wave
and the
synapse.

An action potential is a wave of voltage. Synapses are chemistry between waves. Together: a nervous system.

16Sub-topics
47Key terms
SL+HLLevel
CellsLevel of organisation
C2.2
Why this topic

What this topic answers.

Every sub-topic below feeds at least one of these questions.

Guiding question 1

How are electrical signals generated and moved within neurons?

Guiding question 2

How can neurons interact with other cells?

C2.2.1 – C2.2.7 · Standard Level

7 things to lock in.

The required syllabus content for C2.2, in order. Each card is one lesson-sized checkpoint.

C2.2.1

Neurons as cells within the nervous system that carry electrical impulses

An axon is a long single fibre. Dendrites are multiple shorter fibres. Electrical impulses are conducted along these fibres.

C2.2.2

Generation of the resting potential by pumping to establish and maintain concentration gradients of sodium and potassium ions

They should understand the concept of a membrane polarization and a membrane potential and also reasons that the resting potential is negative.

C2.2.3

Nerve impulses as action potentials that are propagated along nerve fibres

Nerve impulses as action potentials that are propagated along nerve fibres

C2.2.4

Variation in the speed of nerve impulses

Variation in the speed of nerve impulses

C2.2.5

Synapses as junctions between neurons and between neurons and effector cells

Synapses as junctions between neurons and between neurons and effector cells

C2.2.6

Release of neurotransmitters from a presynaptic membrane

Release of neurotransmitters from a presynaptic membrane

C2.2.7

Generation of an excitatory postsynaptic potential

Generation of an excitatory postsynaptic potential

C2.2.1 · Neuron structure

Cells built for signalling.

Neurons have a cell body for the genetic and metabolic essentials, plus long fibres for sending and receiving signals.

Sensory

Receptor → CNS

Carry impulses from sense organs (eyes, ears, skin, etc.) to the central nervous system.

Interneuron

CNS → CNS

Live entirely within the CNS, connecting other neurons. The vast majority of neurons in the brain are interneurons.

Motor

CNS → effector

Carry impulses from the CNS to muscles or glands, triggering action.

Three parts of every neuron:

C2.2.2 · Resting potential

−70 mV, maintained by ATP.

At rest, the inside of a neuron is more negative than the outside. The Na⁺/K⁺ pump establishes this gradient — and uses ATP doing so.

The Na⁺/K⁺ pump moves 3 Na⁺ out and 2 K⁺ in per cycle, using 1 ATP. Combined with large negative organic ions trapped inside, this produces a net negative interior at about −70 mV.

This resting potential is the platform from which action potentials are launched. Maintaining it costs substantial energy — neurons are some of the most metabolically active cells in the body.

C2.2.3 · Nerve impulse

A wave of voltage running down an axon.

A nerve impulse is an action potential propagated along a nerve fibre. It involves the flow of positively charged ions across the membrane.

The signal isn't a chemical or a pulse of light — it's an electrical wave caused by Na⁺ and K⁺ briefly flowing in and out of the axon. Because Na⁺ and K⁺ are charged, their movement changes the membrane potential — and that change propagates as a wave along the fibre.

C2.2.4 · Speed of impulses

Two ways to speed it up.

Conduction speed varies dramatically — from 0.5 m/s in small unmyelinated fibres to 150 m/s in fat myelinated ones.

Wider axon

Squid giant axon

Axon diameter ↑ → conduction speed ↑ (positive correlation). The squid giant axon is up to 1 mm wide and conducts at ~25 m/s — making it indispensable to neuroscience pioneers who needed something big enough to record from.

Myelin

Saltatory conduction

Myelin sheaths insulate most of the axon, leaving small unmyelinated gaps called nodes of Ranvier. Action potentials jump from node to node. Speeds up to 150 m/s. Much more energy-efficient too.

📊

Correlation analysis

Bigger animals tend to have slower nerve impulses if you don't correct for adaptations (negative correlation). But within an animal, larger axons and myelinated axons conduct faster (positive correlation with diameter, with myelination). Use correlation coefficients to quantify strength of these relationships, and R² to assess how much one variable explains the other.

C2.2.5 · Synapses

Chemical bridges between cells.

Where two neurons meet — or a neuron meets a muscle or gland — the signal jumps across a tiny gap chemically. That's a synapse.

A typical (chemical) synapse has three parts:

Signals pass one way only across a synapse — vesicles are on the presynaptic side, receptors on the postsynaptic side. This directional asymmetry gives the nervous system its one-way circuits.

C2.2.6 / C2.2.7 · Synaptic transmission

The handover in five steps.

Action potential arrives → Ca²⁺ enters → vesicles fuse → neurotransmitter diffuses → postsynaptic channels open → next action potential triggered.

  1. Action potential arrives at the axon terminal.
  2. Voltage-gated Ca²⁺ channels open. Ca²⁺ diffuses into the axon terminal.
  3. Ca²⁺ triggers vesicles of neurotransmitter to move to the presynaptic membrane.
  4. Exocytosis. Vesicles fuse with the presynaptic membrane; neurotransmitter is released into the synaptic cleft.
  5. Neurotransmitter diffuses across the cleft and binds receptors on the postsynaptic membrane. At an excitatory synapse (e.g. acetylcholine at NMJ), this opens ligand-gated Na⁺ channels → Na⁺ enters → postsynaptic membrane depolarises → if threshold is reached, an action potential is generated in the postsynaptic cell.

Acetylcholine at the neuromuscular junction

Released by motor neuron axon terminals; binds nicotinic acetylcholine receptors on the muscle fibre's sarcolemma; opens Na⁺ channels; muscle fibre depolarises and contracts. The named example in the IB.

HL extension

Higher Level only.

An extra 9 sub-topics for HL — same syllabus, deeper mechanism.

HL only

Depolarization and repolarization during action potentials

Depolarization and repolarization during action potentials

HL only

Propagation of an action potential along a nerve fibre/axon as a result of local currents

Propagation of an action potential along a nerve fibre/axon as a result of local currents

HL only

Oscilloscope traces showing resting potentials and action potentials

Oscilloscope traces showing resting potentials and action potentials

HL only

Saltatory conduction in myelinated fibres to achieve faster impulses

Saltatory conduction in myelinated fibres to achieve faster impulses

HL only

Saltatory conduction in myelinated fibres to achieve faster impulses

Saltatory conduction in myelinated fibres to achieve faster impulses

HL only

Inhibitory neurotransmitters and generation of inhibitory postsynaptic potentials

Inhibitory neurotransmitters and generation of inhibitory postsynaptic potentials

HL only

Summation of the effects of excitatory and inhibitory neurotransmitters in a postsynaptic neuron

Summation of the effects of excitatory and inhibitory neurotransmitters in a postsynaptic neuron

HL only

Perception of pain by neurons with free nerve endings in the skin

Perception of pain by neurons with free nerve endings in the skin

HL only

Consciousness as a property that emerges from the interaction of individual neurons in the brain

Consciousness as a property that emerges from the interaction of individual neurons in the brain

C2.2.8 · Depolarisation & repolarisation

The action potential in detail.

An action potential is a tight sequence of voltage-gated channel openings and closings — Na⁺ in, then K⁺ out — that produces a sharp spike and recovery.

  1. Resting potential (−70 mV). Na⁺/K⁺ pump maintains gradients.
  2. Stimulus. If depolarisation reaches the threshold potential (~ −55 mV), voltage-gated Na⁺ channels open.
  3. Depolarisation. Na⁺ floods in; membrane potential shoots to +30 mV.
  4. Repolarisation. At +30 mV, Na⁺ channels close, voltage-gated K⁺ channels open. K⁺ flows out; potential drops.
  5. Hyperpolarisation. K⁺ channels close slowly; potential overshoots to below −70 mV.
  6. Return to resting potential. Na⁺/K⁺ pump restores ion gradients. Ready for the next impulse.
C2.2.9 · Propagation

Local currents spread the signal.

When Na⁺ floods into one patch of axon, it diffuses sideways and triggers the next patch's voltage-gated channels — propagating the action potential along the fibre.

The Na⁺ ions don't stay still after entering. They diffuse along the inside of the axon, creating a local current. That current depolarises the adjacent patch of membrane to threshold, opening its Na⁺ channels — propagating the action potential. Behind the wave, the previous patch is repolarising.

C2.2.10 · Oscilloscope traces

What an action potential looks like on a screen.

Plotting membrane potential vs time produces the classic spike — used in the original Hodgkin and Huxley experiments to crack the mechanism.

Phases visible on the trace:

  • Resting potential — flat at −70 mV.
  • Depolarisation — rapid rise to +30 mV.
  • Repolarisation — rapid fall back toward resting.
  • Hyperpolarisation — brief dip below resting potential.
  • Return to resting — flat at −70 mV again.

Total duration: ~3.5 ms. Frequency of impulses can be measured by counting spikes per second.

C2.2.11 · Saltatory conduction

The signal jumps the gaps.

In myelinated axons, action potentials skip from one node of Ranvier to the next — faster and more energy-efficient than continuous conduction.

Myelin (made by Schwann cells in the PNS, oligodendrocytes in the CNS) insulates most of the axon. Sodium-potassium pumps and voltage-gated channels are concentrated at nodes of Ranvier — short gaps between myelin segments.

Action potentials can only occur at the nodes. The signal effectively "jumps" from node to node — saltatory conduction (saltare = to leap). Much faster and uses far less ATP because only the nodes need to actively pump ions.

C2.2.12 · Exogenous chemicals affecting synaptic transmission

Drugs and toxins hack synapses.

External chemicals can interfere with synaptic transmission. Two named examples — one a pesticide, one a recreational drug.

Neonicotinoids

Block insect acetylcholine receptors

Similar shape to acetylcholine. Bind irreversibly to acetylcholine receptors in insects → block synaptic transmission → paralysis and death. Human receptors have a different shape → much less toxicity to humans. But they harm non-target insects like bees.

Cocaine

Blocks dopamine reuptake

Dopamine transporters normally clear dopamine from synaptic clefts. Cocaine blocks these transporters. Dopamine builds up in synapses → continuous stimulation of reward-pathway neurons → euphoria → addiction.

C2.2.13 / C2.2.14 · Inhibitory transmitters & summation

Postsynaptic neurons integrate inputs.

Excitatory inputs push toward firing; inhibitory inputs push away. The neuron adds them all up — and either crosses threshold (fires) or doesn't.

Inhibitory neurotransmitters

Open channels for negative ions (Cl⁻) in the postsynaptic membrane. The postsynaptic cell becomes hyperpolarised — further from threshold. Action potential less likely. GABA and glycine are the major inhibitory neurotransmitters in the human brain.

Summation

A postsynaptic neuron has many synapses with many different presynaptic neurons. Some release excitatory neurotransmitters; some inhibitory. The postsynaptic neuron sums these inputs. If the net effect crosses the threshold potential, it fires; otherwise it doesn't. All-or-nothing — an action potential either happens or it doesn't.

C2.2.15 · Pain perception

Nociceptors and free nerve endings.

Pain receptors in the skin are free nerve endings with channels that open in response to specific harmful stimuli.

Nociceptor channels open in response to:

  • High temperature — TRPV1 channels open above ~43 °C.
  • Acid — H⁺-activated channels open in damaged tissue.
  • Capsaicin from chilli peppers — also activates TRPV1, which is why hot chilli feels hot.

When channels open, positive ions enter, threshold is reached, an action potential is generated. The impulse travels via sensory neurons to the brain, where pain is perceived.

C2.2.16 · Consciousness

The emergent property with the biggest reputation.

No single neuron is conscious. But the interaction of billions of them produces something that is — the brain's most striking emergent property.

Emergent properties arise when components interact — the whole has properties not present in any part. A single H₂O molecule is not "wet"; only many of them together. A single neuron doesn't think; ~86 billion of them, interacting through trillions of synapses, somehow do.

Consciousness — awareness of self and surroundings — is the IB's named example of an emergent property in biology. How exactly the interaction of neurons produces it is the central question of cognitive neuroscience and philosophy of mind.

HL-only key terms

DepolarizationRepolarizationHyperpolarizationThreshold PotentialVoltage-Gated ChannelsLocal CurrentsOscilloscopeOscilloscope TraceSaltatory ConductionMyelinMyelin SheathMyelinated neuronNodes of RanvierExogenous ChemicalsNeonicotinoidsPesticidesCocaineDopamine TransportersInhibitory NeurotransmittersExcitatory NeurotransmittersNociceptorsConsciousnessEmergent Properties
Vocabulary

24 terms to own.

If you can't define one of these in a sentence, that's where to revise next.

NeuronNerve FiberDendriteAxonAxon TerminalSensory NeuronInterneuronResting PotentialSodium-Potassium PumpNerve ImpulseAction PotentialMyelinMyelinated axonsNon-myelinated axonsCorrelationPositive CorrelationNegative CorrelationCoefficient of Determination (R2)SynapsesPresynaptic MembraneSynaptic CleftPostsynaptic membraneNeurotransmitterAcetylcholine

IB Linking Questions

“In what ways are biological systems regulated?”

“How is the structure of specialized cells related to function?”